Could they use “thumb rules” if said species have thumbs? One that us fleshy puny humans use is the right hand rule where a third axis convention (up) corresponds with counterclockwise movement by the hand curl. Of course you could easily use the other hand since they’re arbitrary anyway.
I have a Moonlander split keyboard that is programmable and uses one of these little things for its reset button. My only gripe with it is that they could have designed a place for it to slot into the keyboard itself since it’s very easy to lose it if it isn’t otherwise attached to something.
The reset button is used to apply firmware changes. For this keyboard you can customize the keys to pretty much however you like. I have one of the thumb cluster buttons set to Ctrl-C / Ctrl-V for efficiency. It’s great for making changes to personalize your mechanical keyboard experience… but it requires that pin to make changes lol.
I like it since I can assign Enter to one of the left thumb cluster buttons, you normally can’t hit enter with your left hand on a standard keyboard layout.
All of the color photos by Sergei Prokudin-Gorsky are fantastic and dispell the myth that photos back then were all in black and white. And the developed color film looks so realistic too.